Dr Richard Barnes
Honorary Associate Professor
School of Biological Sciences
Researcher biography
Dr Richard Barnes was for 40 years a staff member of the Department of Zoology of the University of Cambridge, UK, and in 'retirement' he remains attached to that Department and an Emeritus Fellow of St Catharine's College Cambridge. In addition, however, for 3 months of each year he functions as an honorary member of staff of the University of Queensland and of Queensland Museum's Biodiversity Program (October - December inclusive), and likewise (January - March inclusive) of the Department of Zoology of Rhodes University, South Africa.
Book
Barnes, R. S.K. and Hughes, R. N. (2009). An Introduction to Marine Ecology: Third Edition. wiley. doi: 10.1002/9781444313284
Book Chapters
Maxwell, Paul, Connolly, Rod, Roelfsema, Chris, Burfeind, Dana, Udy, James, O'Brien, Kate, Saunders, Megan, Barnes, Richard, Olds, Andrew, Hendersen, Chris and Gilby, Ben (2019). The seagrasses of Moreton Bay Quandamooka: Diversity, ecology and resilience. Moreton Bay Quandamooka and catchment: past, present and future. (pp. 279-298) edited by Ian R. Tibbetts, Peter C. Rothlisberg, David T. Neil, Tamara A. Homburg, David T. Brewer and Angela H. Arthington. Brisbane, QLD, Australia: Moreton Bay Foundation.
York, Paul H., Hyndes, Glenn A., Bishop, Melanie J. and Barnes, Richard S.K. (2018). Faunal assemblages of seagrass ecosystems. Seagrasses of Australia: structure, ecology and conservation. (pp. 541-588) edited by Anthony W. D. Larkum, Gary A. Kendrick and Peter J. Ralph. Cham, Switzerland: Springer. doi: 10.1007/978-3-319-71354-0_17
Barnes, R. S.K. (2001). Lagoons. Encyclopedia of Ocean Sciences. (pp. 377-388) Elsevier Ltd. doi: 10.1016/B978-012374473-9.00091-6
Journal Articles
Barnes, R.S.K. (2023). Seagrass macrobenthic biodiversity does not vary in conformity with a leaky-lagoonal confinement gradient. Marine Environmental Research, 185 105897, 105897. doi: 10.1016/j.marenvres.2023.105897
Barnes, R.S.K. (2022). Interspecific abundance-occupancy relations along estuarine gradients. Marine Environmental Research, 181 105755, 105755. doi: 10.1016/j.marenvres.2022.105755
Barnes, Richard S. K., Claassens, Louw and Seath, Jessica (2022). Where ecologically ‘tis better to go brown than green: enhanced seagrass macrobenthic biodiversity within the canals of a brownfield coastal marina. Biodiversity and Conservation, 31 (12), 2981-2997. doi: 10.1007/s10531-022-02468-9
Barnes, R. S. K. (2021). Patterns of seagrass macrobenthic biodiversity in the warm-temperate Knysna estuarine bay, Western Cape: a review. Aquatic Ecology, 55 (2), 327-345. doi: 10.1007/s10452-021-09848-3
Barnes, R.S.K. (2021). Interspecific relationship of patchiness to occupancy and abundance, as exemplified by seagrass macrobenthos. Ecological Indicators, 121 107083. doi: 10.1016/j.ecolind.2020.107083
Barnes, R. S. K. (2021). Within-species relationship of patchiness to both abundance and occupancy, as exemplified by seagrass macrobenthos. Oecologia, 196 (4), 1107-1117. doi: 10.1007/s00442-021-04985-w
Barnes, R. S.K. (2020). Do species display characteristic intraspecific levels of patchiness in a given habitat type?: the case of intertidal seagrass macrobenthos. Marine Biology, 167 (12) 177. doi: 10.1007/s00227-020-03793-9
Barnes, R. S. K. (2020). Morisita's index of patchiness and edge effects: A response to Butturi-Gomes and Petrere (2020). Ecological Indicators, 117 106648, 1-2. doi: 10.1016/j.ecolind.2020.106648
Barnes, R. S. K. (2020). Do different sympatric seagrasses support macrobenthic faunas of differing composition, abundance, biodiversity or patchiness? Macrobenthos in sympatric seagrasses. Marine Environmental Research, 160 104983, 104983. doi: 10.1016/j.marenvres.2020.104983
Barnes, R.S.K. (2019). Abundance/occupancy/patchiness relations in estuarine seagrass macrobenthos. Estuarine, Coastal and Shelf Science, 228 106360, 106360. doi: 10.1016/j.ecss.2019.106360
Barnes, R. S. K. and Hamylton, Sarah M. (2019). Isometric scaling of faunal patchiness: Seagrass macrobenthic abundance across small spatial scales. Marine Environmental Research, 146, 89-100. doi: 10.1016/j.marenvres.2019.03.011
Myers, A.A., Lowry, J.K. and Barnes, R.S.K. (2018). First record of the genus Eriopisella Chevreux, 1920 (Crustacea, Amphipoda, Senticaudata, Eriopisidae) from Australia, with the description of a new species, Eriopisella moretoni sp. nov. Zootaxa, 4514 (2), 256-262. doi: 10.11646/zootaxa.4514.2.8
Barnes, R. S. K. (2018). Spatial structure of a multi-species guild: the dominant biofilm-grazing microgastropods of seagrass. Hydrobiologia, 827 (1), 293-307. doi: 10.1007/s10750-018-3781-y
Barnes, R. S. K. and Laurie, H (2018). Seagrass macrofaunal abundance shows both multifractality and scale-invariant patchiness. Marine Environmental Research, 138, 84-95. doi: 10.1016/j.marenvres.2018.04.009
Barnes, R.S.K. (2017). Little-known and phylogenetically obscure South African estuarine microgastropods (Mollusca: Truncatelloidea) as living animals. Journal of Natural History, 52 (1-2), 87-113. doi: 10.1080/00222933.2017.1408867
Barnes, R. S. K. (2017). Patterns of benthic invertebrate biodiversity in intertidal seagrass in Moreton Bay, Queensland. Regional Studies in Marine Science, 15, 17-25. doi: 10.1016/j.rsma.2017.07.003
Barnes, R. S. K. (2017). Are seaward pneumatophore fringes transitional between mangrove and lower-shore system compartments?. Marine Environmental Research, 125, 99-109. doi: 10.1016/j.marenvres.2017.01.008
Barnes, R. S. K. (2016). Spatial homogeneity of benthic macrofaunal biodiversity across small spatial scales. Marine Environmental Research, 122, 148-157. doi: 10.1016/j.marenvres.2016.10.006
Barnes, R. S. K. and Hamylton, S. (2016). On the very edge: faunal and functional responses to the interface between benthic seagrass and unvegetated sand assemblages. Marine Ecology Progress Series, 553, 33-48. doi: 10.3354/meps11800
Barnes, R. S.K. and Hendy, I. W. (2015). Seagrass-associated macrobenthic functional diversity and functional structure along an estuarine gradient. Estuarine, Coastal and Shelf Science, 164, 233-243. doi: 10.1016/j.ecss.2015.07.050
Michie, Laura A., Cragg, Simon M., Barnes, R. S.K. and Bennett, Wayne A. (2015). Uca cryptica Naderloo, Türkay & Chen, 2010 (Crustacea: Brachyura: Ocypodidae) is no longer cryptic. Zootaxa, 3981 (2), 291-295. doi: 10.11646/zootaxa.3981.2.10
Barnes, R.S.K. and Hendy, Ian W (2015). Functional uniformity underlies the common spatial structure of macrofaunal assemblages in intertidal seagrass beds. Biological Journal of the Linnean Society, 115 (1), 114-126. doi: 10.1111/bij.12483
Barnes, R. S. K. and Hamylton, Sarah (2015). Uniform functional structure across spatial scales in an intertidal benthic assemblage. Marine Environmental Research, 106 (1), 82-91. doi: 10.1016/j.marenvres.2015.03.006
Barnes, R. S.K. and Barnes, M. K.S. (2014). Spatial uniformity of biodiversity is inevitable if the available species are distributed independently of each other. Marine Ecology Progress Series, 516, 263-266. doi: 10.3354/meps11067
Barnes, R. S. K. (2014). Is spatial uniformity of soft-sediment biodiversity widespread and, if so, over what scales?. Marine Ecology Progress Series, 504, 147-158. doi: 10.3354/meps10770
Barnes, R. S.K. and Barnes, M. K.S. (2014). Biodiversity differentials between the numerically-dominant macrobenthos of seagrass and adjacent unvegetated sediment in the absence of sandflat bioturbation. Marine Environmental Research, 99, 34-43. doi: 10.1016/j.marenvres.2014.05.013
Barnes, R. S.K. (2013). Spatial stability of macrobenthic seagrass biodiversity. Marine Ecology Progress Series, 493, 127-139. doi: 10.3354/meps10546
Barnes, R. S. K. and Hamylton, S. (2013). Abrupt transitions between macrobenthic faunal assemblages across seagrass bed margins. Estuarine Coastal and Shelf Science, 131, 213-223. doi: 10.1016/j.ecss.2013.08.007
Barnes, R. S.K. (2013). Distribution patterns of macrobenthic biodiversity in the intertidal seagrass beds of an estuarine system, and their conservation significance. Biodiversity and Conservation, 22 (2), 357-372. doi: 10.1007/s10531-012-0414-z
Barnes, R. S.K. and Ellwood, M. D.F. (2012). Spatial variation in the macrobenthic assemblages of intertidal seagrass along the long axis of an estuary. Estuarine, Coastal and Shelf Science, 112, 173-182. doi: 10.1016/j.ecss.2012.07.013
Barnes, R. S. K. and Barnes, M. K. S. (2012). Shore height and differentials between macrobenthic assemblages in vegetated and unvegetated areas of an intertidal sandflat. Estuarine Coastal and Shelf Science, 106, 112-120. doi: 10.1016/j.ecss.2012.05.011
Clifton, Julian, Etienne, Michelle, Barnes, David K.A., Barnes, Richard S.K., Suggett, David J. and Smith, David J. (2012). Marine conservation policy in Seychelles: Current constraints and prospects for improvement. Marine Policy, 36 (3), 823-831. doi: 10.1016/j.marpol.2011.11.009
Vieira, Sónia, Coelho, Helena, Nolasco, Rita, Serôdio, João, Barnes, Richard S.K. and Queiroga, Henrique (2012). Repeated cycles of immersion and emersion amplify the crawling rhythm of the intertidal gastropod Hydrobia ulvae. Journal of the Marine Biological Association of the United Kingdom, 92 (3), 565-570. doi: 10.1017/S0025315411000853
Smale, Dan A., Barnes, David K.A., Barnes, Richard S.K., Smith, David J. and Suggett, David J. (2012). Spatial variability in the structure of intertidal crab and gastropod assemblages within the Seychelles Archipelago (Indian Ocean). Journal of Sea Research, 69, 8-15. doi: 10.1016/j.seares.2012.01.002
Barnes, R. S. K. and Ellwood, M. D. F. (2012). The critical scale of small-scale spatial variation in ecological patterns and processes in intertidal macrobenthic seagrass assemblages. Estuarine Coastal and Shelf Science, 98, 119-125. doi: 10.1016/j.ecss.2011.12.014
Barnes, R. S.K. and Ellwood, M.D. Farnon (2011). Macrobenthic assemblage structure in a cool-temperate intertidal dwarf eelgrass bed in comparison with those from lower latitudes. Biological Journal of the Linnean Society, 104 (3), 527-540. doi: 10.1111/j.1095-8312.2011.01738.x
Barnes, R. S.K. and Ellwood, M. D.F. (2011). The significance of shore height in intertidal macrobenthic seagrass ecology and conservation. Aquatic Conservation: Marine and Freshwater Ecosystems, 21 (7), 614-624. doi: 10.1002/aqc.1234
Barnes, R. S. K. and Barnes, M. K. S. (2011). Hierarchical scales of spatial variation in the smaller surface and near-surface macrobenthos of a subtropical intertidal seagrass system in Moreton Bay, Queensland. Hydrobiologia, 673 (1), 169-178. doi: 10.1007/s10750-011-0770-9
Barnes, R. S.K. (2010). Spatial variation in abundance and diversity of the smaller surface and near-surface eelgrass-associated intertidal macrobenthos within a warm-temperate estuarine bay in the Garden Route National Park, RSA. Aquatic Conservation: Marine and Freshwater Ecosystems, 20 (7), 762-772. doi: 10.1002/aqc.1152
Vieira, Sónia, Coelho, Helena, Nolasco, Rita, Serôdio, João, Barnes, Richard and Queiroga, Henrique (2010). The circatidal rhythm of the estuarine gastropod Hydrobia ulvae (Gastropoda: Hydrobiidae). Biological Journal of the Linnean Society, 100 (2), 439-450. doi: 10.1111/j.1095-8312.2010.01428.x
Barnes, R. S.K. (2010). Regional and latitudinal variation in the diversity, dominance and abundance of microphagous microgastropods and other benthos in intertidal beds of dwarf eelgrass, Nanozostera spp. Marine Biodiversity, 40 (2), 95-106. doi: 10.1007/s12526-010-0036-1
Barnes, R. S.K. (2010). A remarkable case of fiddler crab, Uca spp., alpha diversity in Wallacea. Hydrobiologia, 637, 249-253. doi: 10.1007/s10750-009-0007-3
Barnes, R. S.K. (2010). A review of the sentinel and allied crabs (Crustacea: Brachyura: Macrophthalmidae), with particular reference to the genus Macrophthalmus. Raffles Bulletin of Zoology, 58 (1), 31-49.
Barnes, David K.A., Barnes, Richard S.K., Smith, David J. and Rothery, Peter (2009). Littoral biodiversity across scales in the Seychelles, Indian Ocean. Marine Biodiversity, 39 (2), 109-119. doi: 10.1007/s12526-009-0010-y
Barnes, R. S.K., Smith, D. J., Barnes, D. K.A. and Gerlach, J. (2008). Variation in the distribution of supralittoral vegetation around an atoll cay: Desroches (Amirante Islands, Seychelles). Atoll Research Bulletin, 565 (565), 1-6. doi: 10.5479/si.00775630.565.1
Barnes, R. S.K. and Davie, Peter J.F. (2008). A new species of the sentinel crab Macrophthalmus (Mareotis) Barnes, 1967 (Crustacea: Brachyura: Macrophthalmidae) from Western Australia. Zootaxa (1807), 63-68. doi: 10.11646/zootaxa.1807.1.4
Barnes, Richard S.K. (2006). Variation in feeding rate of the intertidal mudsnail Hydrobia ulvae in relation to the tidal cycle. Marine Ecology, 27 (2), 154-159. doi: 10.1111/j.1439-0485.2006.00082.x
Schuwerack, P.-Manuela M., Barnes, Richard S. K., Underwood, Graham J. C. and Jones, Peter W. (2006). Gender and species differences in sentinel crabs (Macrophthalmus) feeding on an Indonesian mudflat. Journal of Crustacean Biology, 26 (2), 119-123. doi: 10.1651/C-2612.1
Barnes, R. S.K. (2005). Interspecific competition and rarity in mudsnails: Feeding interactions between and within Hydrobia acuta neglecta and sympatric Hydrobia species. Aquatic Conservation: Marine and Freshwater Ecosystems, 15 (5), 485-493. doi: 10.1002/aqc.677
Barnes, R. S.K. (2004). The distribution and habitat in the Knysna Estuary of the endemic South African mudsnail Hydrobia knysnaensis and the influence of intraspecific competition and ambient salinity on its abundance. African Journal of Aquatic Science, 29 (2), 205-211. doi: 10.2989/16085910409503811
Barnes, R. S.K. (2003). Feeding rates of continually submerged Hydrobia ulvae vary during the daylight hours. Journal of the Marine Biological Association of the United Kingdom, 83 (6), 1273-1275. doi: 10.1017/S0025315403008646
Barnes, R. S.K. (2003). Interactions between benthic molluscs in a Sulawesi mangal, indonesia: The cerithiid mud-creeper Cerithium coralium and potamidid mud-whelks, Terebralia spp. Journal of the Marine Biological Association of the United Kingdom, 83 (3), 483-487. doi: 10.1017/S0025315403007380h
Barnes, R. S.K. (2002). The occurrence and ecology of a marine hydrobiid mudsnail in the southern hemisphere: The Knysna Estuary, South Africa. African Journal of Ecology, 40 (3), 289-294. doi: 10.1046/j.1365-2028.2002.00375.x
Barnes, R. S.K. (2001). Interference competition in the intertidal mud snail Hydrobia ulvae: Egestion rates revisited. Journal of the Marine Biological Association of the United Kingdom, 81 (3), 491-495. doi: 10.1017/s0025315401004131
Barnes, R. S.K. and De Villiers, C. J. (2000). Animal abundance and food availability in coastal lagoons and intertidal marine sediments. Journal of the Marine Biological Association of the United Kingdom, 80 (2), 193-202. doi: 10.1017/S0025315499001782
Barnes, R. S.K. (1999). What determines the distribution of coastal hydrobiid mudsnails within north-western Europe?. Marine Ecology, 20 (2), 97-110. doi: 10.1046/j.1439-0485.1999.00069.x
Barnes, R. S.K. (1999). The conservation of brackish-water systems: Priorities for the 21st century. Aquatic Conservation: Marine and Freshwater Ecosystems, 9 (6), 523-527. doi: 10.1002/(sici)1099-0755(199911/12)9:6<523::aid-aqc368>3.0.co;2-a
Barnes, R. S.K. and Gandolfi, Silvia Maria (1998). Is the lagoonal mudsnail hydrobia neglecta rare because of competitively-induced reproductive depression and, if so, what are the implications for its conservation?. Aquatic Conservation: Marine and Freshwater Ecosystems, 8 (6), 737-744. doi: 10.1002/(SICI)1099-0755(1998110)8:6<737::AID-AQC312>3.0.CO;2-M
Barnes, R. S.K. (1998). The effects of movement on population density estimates of mudflat epifauna. Journal of the Marine Biological Association of the United Kingdom, 78 (2), 377-385. doi: 10.1017/S0025315400041515
Barnes, R. S.K. (1996). Breeding, recruitment and survival in a mixed intertidal population of the mudsnails Hydrobia ulvae and H. neglecta. Journal of the Marine Biological Association of the United Kingdom, 76 (4), 1003-1012. doi: 10.1017/s0025315400040935
Barnes, R. S.K. (1994). Macrofaunal community structure and life histories in coastal lagoons. Coastal lagoon processes, 60 (C), 311-362. doi: 10.1016/S0422-9894(08)70016-3
Barnes, R. S.K. (1994). Investment in eggs in lagoonal hydrobia ventrosa and life-history strategies in north-west european hydrobia species. Journal of the Marine Biological Association of the United Kingdom, 74 (3), 637-650. doi: 10.1017/S0025315400047718
Barnes, R. S.K. (1994). A Critical Appraisal of the Application of Guélorget and Perthuisot's Concepts of the Paralic Ecosystem and Confinement to Macrotidal Europe. Estuarine, Coastal and Shelf Science, 38 (1), 41-48. doi: 10.1006/ecss.1994.1003
Barnes, R. S.K. (1993). On the nature of the coastal lagoon winkles attributed to Littorina tenebrosa and Littorina saxatilis. CAH.BIOL.MAR., 34 (4), 477-495.
Barnes, R. S.K. (1993). Life-history strategies in contrasting populations of the coastal gastropod Hydrobia. III. Lagoonal versus intertidal-marine H. neglecta. Vie et Milieu, 43 (2-3), 73-83.
Barnes, R. S.K. (1991). European estuaries and lagoons: A personal overview of problems and possibilities for conservation and management. Aquatic Conservation: Marine and Freshwater Ecosystems, 1 (1), 79-87. doi: 10.1002/aqc.3270010107
Barnes, R. S.K. (1991). Dilemmas in the theory and practice of biological conservation as exemplified by British coastal lagoons. Biological Conservation, 55 (3), 315-328. doi: 10.1016/0006-3207(91)90035-8
Barnes, R. S.K. (1991). Two-layered awakening. Nature, 350 (6319). doi: 10.1038/350561a0
Barnes, R. S.K. (1990). Reproductive strategies in contrasting populations of the coastal gastropod Hydrobia ulvae. II. Longevity and life-time egg production. Journal of Experimental Marine Biology and Ecology, 138 (3), 183-200. doi: 10.1016/0022-0981(90)90166-A
Barnes, R. S.K. (1989). The coastal lagoons of Britain: An overview and conservation appraisal. Biological Conservation, 49 (4), 295-313. doi: 10.1016/0006-3207(89)90049-9
Barnes, R. S.K. (1989). What, if anything, is a brackish-water fauna?. Transactions of the Royal Society of Edinburgh: Earth Sciences, 80 (3-4), 235-240. doi: 10.1017/S0263593300028674
Barnes, R. S.K. (1988). The faunas of land-locked lagoons: Chance differences and the problems of dispersal. Estuarine, Coastal and Shelf Science, 26 (3), 309-318. doi: 10.1016/0272-7714(88)90067-4
Barnes, R. S.K. (1988). On reproductive strategies in adjacent lagoonal and intertidal-marine populations of the gastropod hydrobia ulvae. Journal of the Marine Biological Association of the United Kingdom, 68 (3), 365-375. doi: 10.1017/S0025315400043265
Barnes, R. S.K. (1987). Coastal lagoons of East Anglia, U.K. Journal of Coastal Research, 3 (4), 417-427.
Barnes, R. S.K. (1986). Daily activity rhythms in the intertidal gastropod Hydrobia ulvae (Pennant). Estuarine, Coastal and Shelf Science, 22 (3), 325-334. doi: 10.1016/0272-7714(86)90046-6
Barnes, R. S.K. (1981). An experimental study of the pattern and significance of the climbing behaviour of hydrobia ulvae. Journal of the Marine Biological Association of the United Kingdom, 61 (2), 285-299. doi: 10.1017/S0025315400046944
Barnes, R. S.K. (1981). Factors affecting climbing in the coastal gastropod hydrobia ulvae. Journal of the Marine Biological Association of the United Kingdom, 61 (2), 301-306. doi: 10.1017/S0025315400046956
Barnes, R. S.K. (1979). Intrapopulation variation in Hydrobia sediment preferences. Estuarine and Coastal Marine Science, 9 (2), 231-234. doi: 10.1016/0302-3524(79)90117-8
Crawford, R. M., Dorey, A. E., Little, Colin and Barnes, R. S.K. (1979). Ecology of Swanpool, Falmouth. V. Phytoplankton and nutrients. Estuarine and Coastal Marine Science, 9 (2). doi: 10.1016/0302-3524(79)90110-5
Barnes, R. S.K. and Greenwood, J. G. (1978). The response of the intertidal gastropod Hydrobia ulvae (Pennant) to sediments of differing particle size. Journal of Experimental Marine Biology and Ecology, 31 (1), 43-54. doi: 10.1016/0022-0981(78)90135-1
Barnes, R. S.K. (1978). Variation in paragnath number of Nereis diversicolor in relation to sediment type and salinity regime. Estuarine and Coastal Marine Science, 6 (3), 275-283. doi: 10.1016/0302-3524(78)90016-6
Barnes, R. S.K. and Head, S. M. (1977). Variation in paragnath number in some British populations of the estuarine polychaete Nereis diversicolor. Estuarine and Coastal Marine Science, 5 (6), 771-781. doi: 10.1016/0302-3524(77)90048-2
Barnes, R. S.K. (1977). Concluding contribution towards a revision of, and a key to, the genus Macrophthalmus (Crustacea: Brachyura). Journal of Zoology, 182 (2), 267-279. doi: 10.1111/j.1469-7998.1977.tb04159.x
Barnes, R. S.K., Sattelle, D. B., Everton, I. J., Nicholas, W. and Scott, D. H. (1976). Intertidal sands and interstitial fauna associated with different stages of salt-marsh development. Estuarine and Coastal Marine Science, 4 (5). doi: 10.1016/0302-3524(76)90025-6
Barnes, R. S.K. (1973). A Redescription of Macrophthalmus Latipes Borradaile, 1903: An Ocypodid Crab With Portunid-Like Paddles (Decapoda, Brachyura). Crustaceana, 25 (3), 292-296. doi: 10.1163/156854073X00281
Dorey, A. E., Little, Colin and Barnes, R. S.K. (1973). An ecological study of the Swanpool, Falmouth. II. Hydrography and its relation to animal distributions. Estuarine and Coastal Marine Science, 1 (2). doi: 10.1016/0302-3524(73)90067-4
Barnes R.S.K. (1968). Gradual acclimation effects in Macrophthalmus setosus M.Edw. (brachyura: ocypodidae). Life Sciences, 7 (4), 171-173. doi: 10.1016/0024-3205(68)90301-9
Barnes R.S.K. (1968). Individual variation in osmotic pressure of an ocypodid crab. Comparative Biochemistry And Physiology, 27 (2), 447-450. doi: 10.1016/0010-406X(68)90243-0
BARNES, R. S.K. (1968). Aspects of the Australasian zoogeography of the Macrophthalminae (Brachyura, Ocypodidae). Proceedings of the Linnean Society of London, 179 (1), 67-75. doi: 10.1111/j.1095-8312.1968.tb01101.x
Barnes R.S.K. (1968). Relative carapace and chela Proportions in some Ocypodid crabs (Brachyura, Ocypodidae). Crustaceana, 14 (2), 131-136. doi: 10.1163/156854068X00494